These are very similar to plants in the Marchantiales. They are generally less complex in their vegetative anatomy than Marchantia
and its immediate relatives.
They are not very diverse as there are approximately 20 genera and 500
species.
Habitat = Very Wet
Their Growth Form is
simple & Thallose.
They may be Unbranched or have Dichotomous Branching.
Branches may also emerge from the Midrib.This is
sometimes irregular in appearance.
Key Anatomical Features
Lack Air Chambers
This thallus of Monoclea resembles a Green Alga (Chlorophyta)
Pelliaepiphylla Thalli
Lack Scales
Have Rhizoids
Gametangia = Sessile
The Seta elevates the Capsule above the
Thallus.
The Pelliaceae & Pallaviciniaceae are
the two most important families in this Order
An Apical Cell constitutes
the Apical Meristem & thus have Apical Growth.
Dichotomous Branching can occur at the Apex.
The Ventral Mid Rib can
also produce Branches
Thallus
The
Thallus is Smooth & Translucent.
Unlike
Marchantia there is Little Tissue Differentiation.
All Cells are Chlorophyllous and they superficially
resemble Chlorophyta.
There is not the level of tissue differentiation seen with Marchantia.
A Thickened "Midrib"
is present. Rhizoids are produced from this region on the
Ventral surface.
The Midrib contains Xylem-Like
Tissue!
Rhizoids are produced on the ventral side of the Midrib