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Basic Leaf Terminology
Monocots & Ferns

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LeafSheath200.jpg (12643 bytes)
Base of a grass leaf showing the Leaf Sheath which is analogous to a petiole.

l66lily.jpg (38746 bytes)
Lily with Parallel (Longitudinal Striate) Venation

Orch200.jpg (20289 bytes)
Terrestrial Orchid with Parallel (Longitudinal Striate) Venation

TiLeafRedScanWholeAdaxial.jpg (158483 bytes)      TiAdLeafRedScanPart600Crop.jpg (227838 bytes)
Upper (Adaxial) Surface of a Red ki Leaf
TiLeafRedScanAbPart500Sharp.jpg (511653 bytes)
Abaxial Surface

TiLeafGreenScanPart500Crop.jpg (266961 bytes)
Adaxial Surface of a Green ki Leaf
tilef-1-300Crop.jpg (35135 bytes)     ParallelTiClose.jpg (28416 bytes)
ki (Cordyline terminalis) Leaf with Parallel
(Longitudinal Striate) Venation

comisuralBund.jpg (11739 bytes)
Commisural Bundles in a ki leaf link the longitudinal bundles.

TravelerPalm300.jpg (43159 bytes)   Strelitzia.jpg (67789 bytes)      ParVeinWhoMont.jpg (108152 bytes)
Travelers' Palm
is a monocot with large Simple Leaves. Its  partitions are caused by the wind. Much the same occurs with banana (Musa)
. Both have Longitudinal Striate Venation like that shown by the leaf of Strelitzia (Bird of Paradise). The cross-hatch pattern is caused by Commisural Bundles.

Commisural Bundles are very abundant & prominent in the leaf of Maranta (Prayer Plant)
ComisComposLab.jpg (28467 bytes)

PrayerPlant300.jpg (40632 bytes)

Monocots have a Network of Veins!

The prayer plant (Maranta), ki & Traveler's Palm exhibit several vein traits which point out the limitations of the generalization that Monocots have only Parallel Venation and that their venation does not form a Network like Reticulate Venation.

All of the above have Commisural Bundles which results in a close-knit pattern of veins that rivals the contact between veins and the mesophyll seen with Reticulate Venation.

Furthermore, ki and Maranta have apparent "Midribs" and there are "lateral" veins which diverge from them in a pinnate-like pattern (Maranta). These midribs may not contain extremely enlarged veins but they contain many large parallel veins.

Finally, the veins converge and join near the leaf margin forming a closed, integrated vascular system.

The pattern seen in Maranta is called Pinnate-Striate.

TIMribLab300.jpg (38159 bytes)      TiLfMribTBLM500Hue.jpg (332875 bytes)
Cross-sections of ki Midrib

The numerous vascular bundles in the Midrib are evident.

The prominent, fibrous Bundle Sheath is evident in the right-hand image. This provides mechanical strength for the lamina and prevents the disruption of the major veins by physical trauma.

What is the venation of Caladium?
Caladium.jpg (29679 bytes)
Caladium sp. It looks like its Palmate!

Anthurium which is closely related to Caladium and has similar venation.
AnthuriumParadermalVenation400.jpg (55298 bytes)
Note the Reticulate Venation. Is this a Monocot? Yes it is!

Cross-section of Anthurium Leaf Note the Palisade & Spongy Mesophyll which resembles that seen with some Dicots.
AnthuriumX-S400.jpg (34120 bytes)

The Mesophyll of ki leaves is uniform and lacks cellular specializations like those seen above.
TiLfTBLModMod500.jpg (119324 bytes)

Monstera240.jpg (42024 bytes)
Monstera 
LeafAlocasiaAbaxial500.jpg (368665 bytes)
Taro
More monocot leaves that have Pinnate Striate Venation!

The term Striate is preferable to Parallel when Venation in Monocots is considered.

Dichotomous Venation in a Filmy Fern

VeinDichot240.jpg (27907 bytes)    FilmyFernSharp300.jpg (85321 bytes)  
Dichotomous Venation in a Fern
The dark objects are Sori which contain Sporangia.

Dichotomous Venation is characterized by veins which fork at their apex and produce 2 veins of the same size. The branches can develop equally or unequally. The minor veins tend to have the same size rather than getting progressively smaller.

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