Monocot Roots

WB01624_.gif (281 bytes) Root DirectoryWB01624_.gif (281 bytes)

The organization of Monocot roots like Sugarcane is similar to that found in dicots like Ranunculus. However, tracheary elements may be absent from the center of the root.

RotSmilaxWholXS300.jpg (40873 bytes)
Transverse Section of Smilax (Cat Briar)

carrionflowerrootsmilaxlarge.jpg (62355 bytes)
Epifluorescence Image of Smilax Root
CornWholeStele240.jpg (39872 bytes)CornRootStele240.jpg (15120 bytes)

Corn Root: Interface of the Stele & the Cortex-The cells with the largest Diameters are Vessel Members. Follow the Cortical Parenchyma from the top of the image to the first cells which have thick, red-stained walls. These are the Endodermis.

The Endodermis usually develops exceedingly thick secondary walls at levels in the root which are no longer absorbing water. This obscures the Casparian Strips but makes the Endodermis more obvious.

There are usually more xylem arms in monocots.

The number of arms is indicated by the following terms.

Number of Xylem Arms  

Two = Diarch
Three = Triarch
Four = Tetrarch
Five = Pentarch
Six   = Hexarch
More than five = Polyarch

Monocots are usually Polyarch.

Bundles of Phloem alternate with the Xylem arms. The xylem contains many lignified cells in older portions of the root. The phloem stands out because its cells have thin, unlignified cell walls.

ClintoniaRootX-S240.jpg (74675 bytes)

Cross-section of Clintonia (above) & Smilax (below) Roots. Count the number of xylem arms. Both are Monocots

RotSmilaxWholXS300.jpg (40873 bytes)

RotSmilaxEpi-Xylm300.jpg (19191 bytes)

Cross-section of Smilax Root. The tracheary elements of the Xylem are the cells with the largest diameters. Count the number of xylem arms. The Endodermis is the densely stained, lignified layer which surrounds the Stele.

SmilaxRotMaturePhloemXylem300.jpg (41929 bytes)Smilax Stele showing bundles of Phloem  alternating with Xylem Vessel Members.

The cell pattern is distorted by the extreme enlargement of the  Vessel Members.

Smilax root at higher magnification. Locate the Xylem and Phloem as well as the Endodermis, Cortex and Epidermis.


Smilax root showing the Stele, Endodermis, Cortex, SmilaX-sEpiCortCasp300.jpg (51266 bytes)Exodermis & Epidermis. The Exodermis has the same anatomy as the Endodermis and probably restricts water loss to the outside in older roots.

PineappleRootSteleTBl.jpg (217873 bytes)
Inner Cortex and Stele of a Pineapple Root Stained with Toluidine Blue
PineappleRootSteleTBlLite.jpg (185514 bytes)
Stele from Pineapple Root lightly Stained with Toluidine Blue
Locate the Endodermis and Xylem!
orchidaerialrootlargeNik.jpg (72900 bytes)
Autofluorescence Image of an Orchid Aerial Root: Locate the Velamen and the Vascular Cylinder.
PiliRootSteleTBKathyBratten.jpg (101300 bytes) PiliStelePhGlucinkathyBratten.jpg (48947 bytes)
Vascular Tisues in the Root of Pili Grass Stained with
Toluidine Blue (Left) & Phloroglucinol (Right). Note the high degree of Lignification.

Top

WB01624_.gif (281 bytes) Root DirectoryWB01624_.gif (281 bytes)