Plants of Hawaii Volcanoes National Park

Scientific Name Acacia koa A. Gray
Family Fabaceae (Pea Family)

Other Names Hawaiian: koa (transl. "warrior"), sometimes koai`a, koai`e, koa`oha
Status Endemic
Distribution & Habitat Often a dominant species, koa can be bound in the dry to wet forests (Wagner, et al. 1990: 641).

Wide ranging and found between 60 - 2,060 m on all the main islands except Ni`ihau and Koho`olawe (Wagner, et al. 1990: 641).

Carlquist (1980: 285) characterizes Acacia koa as "the most conspicuous trees between the low, open dry forest and the web ohia forest," noting that they usually "form a broad band on the lower slopes."

Koa is often found in dense stands in the mesic forests of the Park.  Koa is the dominant tree species in only one rainforest habitat in the Park, the ola`a forest (Stone and Platt 1990: 173). Ordinarily, `ohi`a is the dominant tree species in the rainforest.

The place to see large intact koa stands is along the Mauna Loa Strip road.

Locations in this Guide Kipuka Puaulu, Park Headquarters, Mauna Loa Strip Road, Top of the Strip Road
General Appearance This is one of the large trees in the Park. Look for it high in the canopy.

The stems are relatively straight.

The trees change as they age. Younger individuals have a whitish gray, smooth bark. Older ones are darker, with the bark becoming orange-brown. The texture also gets rougher and more deeply furrowed.
Mauna Loa Strip Road Mauna Loa Strip Road
Some koa get huge.  You can get the idea from the size of the base of this tree.
Even medium sized trees are quite imposing when you can see them at the edge of a clearing. Kipuka Puaulu

Leaves
Young leaves that are found on saplings, and sometime growing from the larger tree's trunk, are true leaf blades. Note that they are bipinnate.

 

Mauna Loa Strip Road
Older "leaves" are called "phyllodes" because they are not leaf blades, but flattened petioles.  Note the characteristic sickle shape.

They have a characteristic gray-green color.

Top of the Mauna Loa Strip Road

Kipuka Puaulu

Kipuka Puaulu

Top of the Strip Road

Juvenile koa trees often show both types of leaves on the same individual.
Kipuka Puaulu Kipuka Puaulu
Look for phyllodes on the forest floor.  This is a good way to spot the presence of a tree even if its phyllodes are difficult to see high in the canopy. Mid-Elevation Woodland
Flowers & Fruits
The flowers are round, cream-colored or pale yellowish heads less than 0.5 inches wide. Kipuka Puaulu

Kipuka Puaulu

Kipuka Puaulu
Kipuka Puaulu Kipuka Puaulu
Seed pods are flattened and range from 3 to 12 inches long and look somewhat like a lima bean pod.

The pods turn brown when mature.

The seeds are flattish and shiny black.

Kipuka Puaulu
Black seeds are found in the pod.  Small seedlings are sometimes seen on the forest floor.
Kipuka Puaulu Kipuka Puaulu
Phenology Koa produces flowers in the winter and spring. Seed pods mature in late summer and fall.

Phenology Notes

Kipuka Puaulu, tree at entrance Flowers just starting. There are also some mature pods. 7/15/00
near Kilauea Military Camp One tree with abundant flowers. 5/6/01
Kipuka Puaulu Tree at parking lot with a few flowers. 5/6/01
Kipuka Puaulu Same tree without flowers. 8/4/01
     
Natural History

Koa is seen as a "keystone" species in the Mauna Loa Strip section of the park. It is host to a number of other species, including insects and birds.

Koa is a prized wood and it has been logged for two centuries. This makes koa a rare tree in the lowland forests. High-elevation lands lost much of their koa forests through their conversion to grazing lands for cattle.

The picture on the right was taken at the Hawaii Volcano Observatory and shows a light-green patch of pasture adjacent to the Park boundary.

Cattle browse the seedlings, suckers and foliage. This has limited the growth and reproduction of koa in areas that are not protected.

Cattle grazing was (finally) eliminated from HVNP in 1948. Goats were excluded in the early 1980s.

Volcano Observatory: Light-green area is pasture land
A fire burned more than 2,000 acres in the area around the Strip Road in 1975. This impacted the koa woodlands, perhaps by stimulating the growth of the trees by stimulating root sucker development.  Therefore, most of the smaller trees shown in these two pictures are about 25 years old.

Look for the presence or evidence of a large individual in the middle of stands of smaller-trunk trees.

As a legume, this species can fix atmospheric nitrogen (Stone and Pratt 1990: 243).  This may contribute to the palatability of the shoots and suckers.  Cows and goats eat these structures if these grazers are allowed into koa stands.  In the 1940s, these animals were prevalent in the Park.  Cattle were removed by 1948 and goats in the 1980s.

Mauna Loa Strip Road

Mauna Loa Strip Road

The clump of koa trees growing at the top of the strip road is at the top of the elevation range for this species.

Note that the trees in this "ring" are about the same size.  It appears that they may all be the result of sucker growth from an older individual that was once growing in the center.

Top of the Mauna Loa Strip Road

 

Conservation Cattle grazing in the Park has had a strong impact on the Acacia koa populations.  This has led some observers, such as Carlquist (1980: 287) to characterize them with a tendency to "form a woodland which is not dense but can be rather open, carpeted by the old phyllodes."  Now that cattle grazing has been stopped in the Park, rather thick stands of trees can be observed. 

This doesn't mean that there are a lot of places where you can see large stands of straight-trunked, old koa.  This is the species that was favored for Hawaiian canoes.  The lowland koa forests were logged and the upland forests converted to cattle pastures even by the end of the 19th century (Stone and Pratt 1990: 242).

This is a fire adapted species (Stone and Pratt 1990: 243).  Fire stimulates growth from the suckers (or shoots) from the shallow roots.  Some of the dense koa forests on the lower portions of the strip road appear to be the result of a 1975 burn that covered some 2,000 acres (Stone and Pratt 1990: 243).

Miscellaneous Photos Kipuka Puaulu (huge tree)
Other Notes Carlquist (1980: 105) has speculated that koa may have dispersed to Hawaii by floating from Australia. Wagner, et al. (1990: 642) relate that this species appears to be closely related to one in the Mascarene Islands in the Indian Ocean.  Further, both species may have a common Australian ancestor.

This species also includes what were once considered separate species, such as A. koaia.


References Carlquist, Sherwin 1980. Hawaii: A Natural History (2nd ed.). Pacific Tropical Botanical Gardens, Lawai, Kauai, Hawaii.

Wagner, Warren, Derral Herbst and S. H. Sohmer 1990. Manual of the Flowering Plants of Hawai`i. University of Hawaii Press.

Links to Other Sites How to plant a native Hawaiian garden (fix this link)

Return to:

 

Last Updated: 08/05/03